As a consequence, Wellingtons were switched to night operations and participated in the first night raid on Berlin on 25 August 1940. In the first 1,000-bomber raid on Cologne, on 30 May 1942, 599 out of 1,046 aircraft were Wellingtons (101 of them were flown by Polish aircrew). With Bomber Command, Wellingtons flew 47,409 operations, dropped 41,823 tons (37,941 tonnes) of bombs and lost 1,332 aircraft in action.
Coastal Command Wellingtons carried out anti-submarine duties and sank their first enemy vessel on 6 July 1942. DWI versions (see below) fitted with a 48 ft (14.63 m) diameter metal hoop were used for exploding enemy mines by generating a powerful magnetic field as it passed over them. In 1944, Wellingtons of Coastal Command were deployed to Greece, and performed various support duties during the RAF interference in the Greek Civil War. A few Wellingtons were operated by the Hellenic Air Force.
While the Wellington was superseded in the European Theatre, it remained in operational service for much of the war in the Middle East and in 1942, Wellingtons based in India became the RAF's first long-range bomber operating in the Far East. It was particularly effective with the South African Air Force in North Africa. This versatile aircraft also served in anti-submarine duties with 26 Squadron SAAF based in Takoradi, Gold Coast (now Ghana).
In late 1944, a radar-equipped Wellington XIV from 407 Sqn. RCAF was modified for use by the RAF's Fighter Interception Unit as what would now be described as an airborne early warning and control aircraft. It operated at an altitude of some 4,000 ft (1,219 m) over the North Sea to control a de Havilland Mosquito and a Bristol Beaufighter fighter intercepting Heinkel He 111 bombers flying from Dutch airbases, and carrying out airborne launches of the V-1 flying bomb. The FIU operator's on the Wellington would search for the He-111 aircraft climbing to launch altitude, then direct the Beaufighter to the bomber, while the Mosquito would attempt to intercept the V-1 if launched.
The Wellington is listed in the appendix to the novel KG 200 as one flown by the German secret operations unit KG 200, which also tested, evaluated and sometimes clandestinely operated captured enemy aircraft during the Second World War.
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